Gay Waluigi and Fat Wario were raised in the Mushroom Kingdom. They eventually got in trouble from Oiraw, who kidnapped Princess Weach and locked her in his 7 Weegee hotels. There, Gay Waluigi found his fascination with cheese strings (as opposed to spaghetti). Once they rescued her, they returned to the Mushroom Kingdom. They met Gay Luigi and Fat Mario, and instantly became enemies.
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